This paper addresses the mathematics of modelling red queen oscillations. Specifically the interactive effective of red queen dynamics, and ecological dynamics which previously has not been addressed fully. They show that the time taken for extinction (the time for a genotype to me lost), is faster when populations size are smaller, competition is on a local scale and variation in population size is taken in to account. This illustrates that larger population although increasing levels of intra-specific competition can help dampen the effects of inter specific competition and red queen dynamics. It also demonstrates the benefit of breeding with populations outside of your immediate core area.
Jack Richardson - Journal update
Link to paper: https://bmcevolbiol.biomedcentral.com/articles/10.1186/s12862-019-1562-5
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